To cross twice in hybridization, as (A × B) × (C × D); for example, in commercial hybrid seed corn, A through D are classically inbreds, and their grandoffspring is the seed for sale.
Origin
First recorded in 1834 from thieves' slang cross (or on the cross) to refer to something dishonest, a play on straight/square, and therefore a crook going back on his partners would be crossing the crossers, or double-crossing.
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