Sense 2 (“joke”) derives from the practice of tapping one’s toes to indicate pleasure; compare toe-tappingly. Sense 3 (“closeted homosexual, especially one who solicits gay sex”) derives from the 2007 Larry Craig scandal in which then-Senator Larry Craig (born 1945) of Idaho, USA, was arrested after tapping his right foot in a men’s restroom, an action seen as covert solicitation of sex.
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