A beetle whose defence mechanism enables it to eject a hot aqueous mixture of chemicals at predators. They come from a number of subfamilies: Carabinae (subfamily) (e.g., Brachinus crepitans (species)), Metriinae (subfamily) (e.g., Stenaptinus insignis (species)) and Paussinae (subfamily) (e.g., Goniotropis nicaraguensis (species)).
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